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The
contents of the egg undergo a series of determinate cleavages
giving rise to the blastula and
triploblastic gastrula stages. Finally, the first stage larva is
formed. The embryo and first-stage larvae are highly resistant to
moisture stress". The first moult occurs within the egg to give rise
to the second-stage larva that is ready to hatch out. The hatching
can occur freely in water or moist soil without any requirement of
root exudates or hatching factors. However, under adverse conditions
of moisture, osmotic pressure, temperature, etc., the hatching is
inhibited or postponed to ensure longer survival. The gelatinous egg
matrix also assists in survival. For instance, due to hygroscopic
nature, it absorbs ambient moisture and the moisture content in the
egg sac remains higher than that in drying soil. Further, under high
degree of moisture stress the high osmotic pressure in the matrix
may inhibit hatching.
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